Professor of Ophthalmology
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| 1975 | Diploma in Biology |
| 1977 | ‘Staatsexamen’ in Medicine |
| 1981 | Medical Thesis |
| 2000- | Director, Institute for Brain Research, Bremen University |
| 1999- | Chair Human Neurobiology, Bremen University |
| 1999-2001 | Visiting Professor, University College, London |
| 1998- | Head of Department, Dept. Optometry & Visual Science, City University London (continued as Professor on part-time basis) |
| 1996-1998 | Board of Directors, Centre for Drug Research, Tübingen |
| 1996 | Wiersma-Professor, Division of Biology, California Institute of Technology |
| 1994-1998 | Professor of Ophthalmology; Head of Section ‘Visual Science’ |
| 1992-1998 | Head, German Research Council Group Neuro-Ophthalmology |
| 1989-1990 | Visiting Scientist, Centre for Biological Information Processing, MIT |
| 1988-1994 | ‘Heisenberg’-Professorship (German Research Council Stipend) |
| 1981-1988 | Head of Electrophysiological Laboratory, University Eye-Hospital, Tübingen; tenured position |
| 1984 | Visiting Scientist, Dept. Anatomy & Physiology, UC Berkeley |
| 1977-1981 | Doctorate Student – since 1980 PostDoc, Max-Planck-Institute for Biological Cybernetics, Tübingen |